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Publications in peer-reviewed journals

2018 & in press

[55] Jablonski D., Gvoždík V., Choleva L., Jandzik D., Moravec J., Mačát Z., Veselý M.: Tracing the maternal origin of the common wall lizard (Podarcis muralis) on the northern range margin in Central Europe. Mitochondrion, in press.

[54] Charles K.L., Bell R.C., Blackburn D.C., Burger M., Fujita M.K., Gvoždík V., Jongsma G.F.M., Talla Kouete M., Leaché A.D., Portik D.M., 2018: Sky, sea, and forest islands: Diversification in the African leaf-folding frog Afrixalus paradorsalis (Anura: Hyperoliidae) of the Lower Guineo-Congolian rain forest. Journal of Biogeography 45: 1781–1794. [pdf]

[53] Dolinay M., Tasse Taboue G.C., Fokam E.B., Gvoždík V., 2018: Geographic and altitudinal range extension of Arthroleptis palava (Anura: Arthroleptidae) from the Cameroon Volcanic Line. Herpetology Notes 11: 557560. [pdf]

[52] Gvoždík V., Dolinay M., Zassi-Boulou A.-G., Badjedjea Babangenge G., 2018: New data on Trachylepis makolowodei from Central Africa. Herpetology Notes 11: 515–518. [pdf]

[51] Mikulíček P., Jablonski D., Páleník M., Gvoždík V., Jandzik D., 2018: Characterization of microsatellite markers in the genera Anguis and Pseudopus (Reptilia: Anguidae). Salamandra 54: 158–162. [pdf]

[50] Kotsakiozi P., Jablonski D., Ilgaz Ç., Kumlutaş Y., Avcı A., Meiri S., Itescu Y., Kukushkin O., Gvoždík V., Scillitani G., Roussos S.A., Jandzik D., Kasapidis P., Lymberakis P., Poulakakis N., 2018: Multilocus phylogeny and coalescent species delimitation in Kotschy’s gecko, Mediodactylus kotschyi: hidden diversity and cryptic species. Molecular Phylogenetics and Evolution 125: 177187. [pdf]

[49] Jandzik D., Jablonski D., Zinenko O., Kukushkin O.V., Moravec J., Gvoždík V., 2018: Pleistocene extinctions and recent expansions in an anguid lizard of the genus Pseudopus. Zoologica Scripta 47: 21–32. [pdf]



[48] Jablonski D., Najbar B., Grochowalska R., Gvoždík V., Strzała T., 2017: Phylogeography and postglacial colonization of Central Europe by Anguis fragilis and Anguis colchica. Amphibia-Reptilia 38: 562–569. [pdf]

[47] Šmíd J., Moravec J., Gvoždík V., Štundl J., Frynta D., Lymberakis P., Kapli P., Wilms T., Schmitz A., Shobrak M., Yousefkhani S.H., Rastegar-Pouyani E., Castilla A.M., Els J., Mayer W., 2017: Cutting the Gordian Knot: Phylogenetic and ecological diversification of the Mesalina brevirostris species complex (Squamata, Lacertidae). Zoologica Scripta 46: 649–664. [pdf]

[46] Bell R.C., Parra J.L., Badjedjea G., Barej M.F., Blackburn D.C., Burger M., Channing A., Dehling J.M., Greenbaum E., Gvoždík V., Kielgast J., Kusamba C., Lötters S., McLaughlin P.J., Nagy Z.T., Rödel M.-O., Portik D.M., Stuart B.L., VanDerWal J., Zassi-Boulou A.-G., Zamudio K.R., 2017: Idiosyncratic responses to climate-driven forest fragmentation and marine incursions in reed frogs from Central Africa and the Gulf of Guinea Islands. Molecular Ecology 26: 5223–5244. [pdf]

[45] Lehr E., Moravec J., Cusi J.C., Gvoždík V., 2017: A new minute species of Pristimantis (Amphibia: Anura: Craugastoridae) with a large head from the Yanachaga-Chemillén National Park in central Peru, with comments on the phylogenetic diversity of Pristimantis occurring in the Cordillera Yanachaga. European Journal of Taxonomy 325: 1–22. [pdf]

[44] Cusi J.C., Moravec J., Lehr E., Gvoždík V., 2017: A new species of semiarboreal toad of the Rhinella festae group (Anura, Bufonidae) from the Cordillera Azul National Park, Peru. ZooKeys 673: 21–47. [pdf]

[43] Doherty-Bone T.M., Gvoždík V., 2017: The Amphibians of Mount Oku, Cameroon: an updated species inventory and conservation review. ZooKeys 643: 109–139. [pdf]

[42] Zimkus B.M., Lawson L.P., Barej M.F., Barratt C.D., Channing A., Dash K.M., Dehling J.M., Du Preez L.; Gehring P.-S., Greenbaum E., Gvoždík V., Harvey J., Kielgast J., Kusamba C., Nagy Z.T., Pabijan M., Penner J., Rödel M.-O., Vences M., Lötters S., 2017: Leapfrogging into new territory: How Mascarene ridged frogs diversified across Africa and Madagascar to maintain their ecological niche. Molecular Phylogenetics and Evolution 106: 254–269. [pdf]

[41] Moravec J., Arista Tuanama I., Gagliardi Urrutia G., Gvoždík V., 2017 “2016”: Amphibians and reptiles recorded in the Conservation Area Imiría in the Ucayali region in Peru. Acta Societatis Zoologicae Bohemicae 80: 317–341. [pdf]



[40] Jablonski D., Jandzik D., Mikulíček P., Džukić G., Ljubisavljević K., Tzankov N., Jelić D., Thanou E., Moravec J., Gvoždík V., 2016: Contrasting evolutionary histories of the legless lizards slow worms (Anguis) shaped by the topography of the Balkan Peninsula. BMC Evolutionary Biology 16: 99. [pdf]

[xx] Ceríaco L.M.P., Gutiérrez E.E., Dubois A. et al. (487 signatories), 2016: Photography-based taxonomy is inadequate, unnecessary, and potentially harmful for biological sciences. Zootaxa 4196 (3): 435–445. [pdf]

[39] Liedtke H.C., Müller H., Rödel M.-O., Menegon M., Gonwouo L.N., Barej M.F., Gvoždík V., Schmitz A., Channing A., Nagel P., Loader S.P., 2016: No ecological opportunity signal on a continental scale? Diversification and life-history evolution of African true toads (Anura: Bufonidae). Evolution 70: 1717–1733. [pdf]








[38] Gvoždík V., Canestrelli D., García-París M., Moravec J., Nascetti G., Recuero E., Teixeira J., Kotlík P., 2015: Speciation history and widespread introgression in the European short-call tree frogs (Hyla arborea sensu lato, H. intermedia and H. sarda). Molecular Phylogenetics and Evolution 83: 143–155.  [pdf]

[37] Evans B.J., Carter T.F., Greenbaum E., Gvoždík V., Kelley D.B., McLaughlin P.J., Pauwels O.S.G., Portik D.M., Stanley E.L., Tinsley R.C., Tobias M.L., Blackburn D.C., 2015: Genetics, morphology, advertisement calls, and historical records distinguish six new polyploid species of African clawed frog (Xenopus, Pipidae) from West and Central Africa. PLoS ONE 10 (12): e0142823.  [pdf]

[36] Furman B.L.S., Bewick A.J., Harrison T.L., Greenbaum E., Gvoždík V., Kusamba C., Evans B.J., 2015: Pan-African phylogeography of a model organism, the African clawed frog ‘Xenopus laevis. Molecular Ecology 24: 909–925.  [pdf]

[35] Bell R.C., Drewes R.C., Channing A., Gvoždík V., Kielgast J., Lötters S., Stuart B.L., Zamudio K.R., 2015: Overseas dispersal of Hyperolius reed frogs from Central Africa to the oceanic islands of São Tomé and Príncipe. Journal of Biogeography 42: 65–75.  [pdf]

[34] Penske S., Gvoždík V., Menegon M., Loader S.P., Müller H., 2015: Description of the tadpole of Leptopelis cf. grandiceps (Amphibia: Anura: Arthroleptidae) from the Uluguru Mountains, Tanzania. Herpetological Journal 25: 61–64.  [pdf]

[33] Šmíd J., Moravec J., Kratochvíl L., Nasher A.K., Mazuch T., Gvoždík V., Carranza S., 2015: Multilocus phylogeny and taxonomic revision of the Hemidactylus robustus species group (Reptilia, Gekkonidae) with descriptions of three new species from Yemen and Ethiopia. Systematics and Biodiversity 13 (4): 346–368.  [pdf]





[32] Gvoždík V., Tillack F., Menegon M., Loader S.P., 2014: The status of Leptopelis barbouri Ahl, 1929 and eleven other nomina of the current tree-frog genus Leptopelis (Arthroleptidae) described from East Africa, with a redescription of Leptopelis grandiceps Ahl, 1929. Zootaxa 3793 (1): 165–187.  [pdf]

[31] Nagy Z.T., Gvoždík V., Meirte D., Collet M., Pauwels O.S.G., 2014: New data on the morphology and distribution of the enigmatic Schouteden’s sun snake, Helophis schoutedeni (de Witte, 1922) from the Congo Basin. Zootaxa 3755 (1): 96–100.  [pdf]

[30] Barej M.F., Rödel M.-O., Loader S.P., Menegon M., Gonwouo N.L., Penner J., Gvoždík V., Günther R., Bell R.C., Nagel P., Schmitz A., 2014: Light shines through the spindrift – Phylogeny of African torrent frogs (Amphibia, Anura, Petropedetidae). Molecular Phylogenetics and Evolution 71: 261–273.  [pdf]

[29] Moravec J., Lehr E., Cusi J.C., Córdova J.H., Gvoždík V., 2014: A new species of the Rhinella margaritifera species group (Anura, Bufonidae) from the montane forest of the Selva Central, Peru. ZooKeys 371: 35–56.  [pdf]


[28] Gvoždík V., Javůrková V., Kopecký O., 2013: First evidence of a paedomorphic population of the smooth newt (Lissotriton vulgaris) in the Czech Republic. Acta Herpetologica 8 (1): 53–57.  [pdf]

[27] Gvoždík V., Benkovský N., Crottini A., Bellati A., Moravec J., Romano A., Sacchi R., Jandzik D., 2013: An ancient lineage of slow worms, genus Anguis (Squamata: Anguidae), survived in the Italian Peninsula. Molecular Phylogenetics and Evolution 69: 1077–1092.  [pdf]

[26] Zimkus B.M., Gvoždík V., 2013: Sky Islands of the Cameroon Volcanic Line: a diversification hotspot for puddle frogs (Phrynobatrachidae: Phrynobatrachus). Zoologica Scripta 42 (6): 591–611.  [pdf]

[25] Jandzik D., Avcı A., Gvoždík V., 2013: Incongruence between taxonomy and genetics: three divergent lineages within two subspecies of the rare Transcaucasian rat snake (Zamenis hohenackeri). Amphibia-Reptilia 34: 579–584.  [pdf]

[24] Šmíd J., Moravec J., Kratochvíl L., Gvoždík V., Nasher A.K., Busais S.M., Wilms T., Shobrak M.Y., Carranza S., 2013: Two newly recognized species of Hemidactylus (Squamata, Gekkonidae) from the Arabian Peninsula and Sinai, Egypt. ZooKeys 355: 79–107.  [pdf]

[23] Šmíd J., Carranza S., Kratochvíl L., Gvoždík V., Nasher A.K., Moravec J., 2013: Out of Arabia: A complex biogeographic history of multiple vicariance and dispersal events in the gecko genus Hemidactylus (Reptilia: Gekkonidae). PLoS ONE 8 (5): e64018.  [pdf]

[22] Šrámek J., Gvoždík V., Benda P., 2013: Hidden diversity in bent-winged bats (Chiroptera: Miniopteridae) of the Western Palaearctic and adjacent regions: implications for taxonomy. Zoological Journal of the Linnean Society 167: 165–190.  [pdf]

[21] Kindler C., Böhme W., Corti C., Gvoždík V., Jablonski D., Jandzik D., Metallinou M., Široký P., Fritz U., 2013: Mitochondrial phylogeography, contact zones and taxonomy of grass snakes (Natrix natrix, N. megalocephala). Zoologica Scripta 42 (5): 458–472.  [pdf]

[20] Jungfer K.-H., Faivovich J., Padial J.M., Castroviejo-Fisher S., Lyra M.M., Berneck B.V.M., Iglesias P.P., Kok P.J.R., MacCulloch R.D., Rodrigues M.T., Verdade V.K., Torres Gastello C.P., Chaparro J.C., Valdujo P.H., Reichle S., Moravec J., Gvoždík V., Gagliardi-Urrutia G., Ernst R., De la Riva I., Means D.B., Lima A.P., Señaris J.C., Wheeler W.C., Haddad C.F.B., 2013: Systematics of spiny-backed treefrogs (Hylidae: Osteocephalus): an Amazonian puzzle. Zoologica Scripta 42 (4): 351–380.  [pdf]

[19] Nagy Z.T., Kusamba C., Collet M., Gvoždík V., 2013: Notes on the herpetofauna of western Bas-Congo, Democratic Republic of the Congo. Herpetology Notes 6: 413–419.  [pdf]


[18] Gvoždík V., Jandzik D., Cordos B., Rehák I., Kotlík P., 2012: A mitochondrial DNA phylogeny of the endangered vipers of the Vipera ursinii complex. Molecular Phylogenetics and Evolution 62: 1019–1024.  [pdf]

[17] Baláž V., Kopecký O., Gvoždík V., 2012: Presence of the amphibian chytrid pathogen confirmed in Cameroon. Herpetological Journal 22: 191–194.  [pdf]

[16] Jablonski D., Jandzik D., Gvoždík V., 2012: New records and zoogeographic classification of amphibians and reptiles from Bosnia and Herzegovina. North-Western Journal of Zoology 8 (2): 324–337.  [pdf]

[15] Gvoždík V., Kopecký O., 2012: Geographic distribution: Phlyctimantis leonardi. Herpetological Review 43 (1): 98.  [pdf]


[14] Moravec J., Kratochvíl L., Amr Z.S., Jandzik D., Šmíd J., Gvoždík V., 2011: High genetic differentiation within the Hemidactylus turcicus complex (Reptilia: Gekkonidae) in the Levant, with comments on the phylogeny and systematics of the genus. Zootaxa 2894: 21–38.  [pdf]

[13] Gvoždík V., 2011: Geographic distribution: Cardioglossa schioetzi. Herpetological Review 42 (1): 106–107.  [pdf]

[12] Gvoždík V., 2011: Geographic distribution: Cardioglossa alsco. Herpetological Review 42 (1): 106.  [pdf]


[11] Gvoždík V., Moravec J., Klütsch C., Kotlík P., 2010: Phylogeography of the Middle Eastern tree frogs (Hyla, Hylidae, Amphibia) as inferred from nuclear and mitochondrial DNA variation, with a description of a new species. Molecular Phylogenetics and Evolution 55: 1146–1166.  [pdf]

[10] Gvoždík V., Jandzik D., Lymberakis P., Jablonski D., Moravec J., 2010: Slow worm, Anguis fragilis (Reptilia: Anguidae) as a species complex: Genetic structure reveals deep divergences. Molecular Phylogenetics and Evolution 55: 460–472.  [pdf]

[9] Blackburn D.C., Gvoždík V., Leaché A.D., 2010: A new squeaker frog (Arthroleptidae: Arthroleptis) from the mountains of Cameroon and Nigeria. Herpetologica 66 (3): 335–348. [& Erratum: Herpetologica 66 (4): 487.]  [pdf]

[8] Benda P., Gvoždík V., 2010: Taxonomy of the genus Otonycteris (Chiroptera: Vespertilionidae: Plecotini) as inferred from morphological and mtDNA data. Acta Chiropterologica 12(1): 83–102.  [pdf]

[7] Barej M.F., Ineich I., Gvoždík V., Lhermitte-Vallarino N., Gonwouo N.L., LeBreton M., Bott U., Schmitz A., 2010: Insights into chameleons of the genus Trioceros (Squamata: Chamaeleonidae) in Cameroon, with the resurrection of Chamaeleon serratus Mertens, 1922. Herpetologia Koenigiana. Bonn zoological Bulletin 57 (2): 211–229.  [pdf]

[6] Gvoždík V., 2010: Second species of tree frog, Hyla orientalis (formely H. arborea), from Iran confirmed by acoustic data. Herpetology Notes 3: 41–44.  [pdf]


[5] Moravec J., Aparicio J., Guerrero-Reinhard M., Calderón G., Jungfer K.-H., Gvoždík V., 2009: A new species of Osteocephalus (Anura: Hylidae) from Amazonian Bolivia: first evidence of tree frog breeding in fruit capsules of the Brazil nut tree. Zootaxa 2215: 37–54.  [pdf]


[4] Gvoždík V., Moravec J. & Kratochvíl L., 2008: Geographic morphological variation in parapatric Western Palearctic tree frogs, Hyla arborea and Hyla savignyi: are related species similarly affected by climatic conditions? Biological Journal of the Linnean Society 95: 539–556.  [pdf]

[3] Gvoždík V., 2008: Geographical distribution: Dipsadoboa underwoodi Rasmussen, 1993 – Underwood’s Rear-fanged Tree-Snake. Cameroon. African Herp News 45: 23–25.  [pdf]


[2] Hořák D., Gvoždík V., Reif J., Klvaňa P., 2007: A nest of Grey-necked Picathartes Picathartes oreas beside a temporal stream. Bulletin of the African Bird Club 14 (2): 183–184.  [pdf]


[1] Gvoždík V., Moravec J., 2005: Variation of Hyla savignyi: A color pattern of Cypriote and mainland populations. In: Ananjeva N. & Tsinenko O. (Eds.): Herpetologia Petropolitana. Proc. of the 12th Ord. Gen. Meeting Soc. Eur. Herpetol., August 12–16, 2003, St. Petersburg. Russian Journal of Herpetology 12 (Suppl.): 32–34.  [pdf]



Book chapters

In: Moravec J. (Ed.), 2015: Fauna ČR. Plazi – Reptilia. Praha: Academia, 531 pp (in Czech).obalka-plazi-def4.indd

[Fauna of the Czech Republic. Reptiles – Reptilia.]

ISBN: 978-80-200-2416-9


Moravec J., Gvoždík V., 2015: Rod Anguis Linnaeus, 1758 – slepýš, p: 236 (in Czech). [Slow worm – Anguis Linnaeus, 1758]

Gvoždík V.
, Moravec J., 2015: Anguis fragilis Linnaeus, 1758 – slepýš křehký, pp. 237–261 (in Czech). [Common European slow worm – Anguis fragilis Linnaeus, 1758]

Moravec J., Gvoždík V., 2015: Anguis colchica (Nordmann, 1840) – slepýš východní, pp. 262–275 (in Czech). [Eastern slow worm – Anguis colchica (Nordmann, 1840)]

Gvoždík V.
, Moravec J., Zavadil V., Jeřábková L., 2015: Slepýš křehký a slepýš východní – výskyt v České republice, pp. 275–278 (in Czech). [Anguis fragilis and A. colchica – distribution in the Czech Republic]

Gvoždík V.
, 2015: Genetická data. Vipera berus (Linnaeus, 1758) – zmije obecná, p. 401 (in Czech). [Genetic data. Common adder – Vipera berus (Linnaeus, 1758)]

Václav Gvoždík, Research on the Evolutionary biology of amphibians & reptiles | Last updated August 2018